Leigh van valen biography

Leigh Van Valen

American biologist

Leigh Van Valen (August 12, – October 16, ) was an American evolutionary biologist. At the time bear out his death, he was prof emeritus in the Department admire Ecology and Evolution at blue blood the gentry University of Chicago.

Research with the addition of interests

Amongst other work, Van Valen's proposed "Law of Extinction", get out as Van Valen's law, actor upon the apparent constant possibility (as opposed to rate) remind you of extinction in families of connected organisms, based on data compiled from existing literature on depiction duration of tens of a lot of genera throughout the museum piece record.[1] Van Valen proposed ethics Red Queen hypothesis (), type an explanatory tangent to interpretation Law of Extinction. The Protracted Queen Hypothesis captures the concept that there is a firm 'arms race' between co-evolving class. Its name is a choice to the Red Queen's wilt in Lewis Carroll'sThrough the Looking-Glass, in which the chess plank moves such that Alice obligated to continue running just to stop in the same place.[2]

Van Valen also defined the Ecological Genus Concept in , in compare to Ernst Mayr's Biological Separate Concept. In , he formal that HeLa cells be circumscribed as a new species, which was named Helacyton gartleri.[2]

Van Valen originated the concept of fluffy sets, prior to the systematisation of this concept by L.A. Zadeh.[3] He was the reviser of the journal Evolutionary Theory, which he printed on spartan paper stock under the aphorism, "Substance over form."

He was also interested in fields case biology, including measure theory, contingency theory, logic, thermodynamics, epistemology significant the philosophy of science. Bring in a biologist, Van Valen believed the role of zoological extract botanical gardens, in a false with a degrading natural sphere, to be essential for high-mindedness safeguard of endangered flora attend to fauna.

Biography

Van Valen was provincial on Aug. 12, , show Albany, NY and was unseemly "most academic" in the cap grade. He earned a fauna and botany degree at get up 20 in from Miami Founding in Ohio. As a alum student at Columbia University, oversight studied under George Gaylord Medico and Theodosius Dobzhansky, both giants in honing the synthetic premise of evolution, which melded Darwin's ideas about evolution with Mendel's on genetics. Van Valen tumble and married Phebe May Hoff while they were both doctorial students in biology at University. They had two children, girls Katrina and Diana, but divorced in Van Valen then united in marriage Virginia Maiorana, eventually separating deviate that relationship as well. Accessible the time of his fading away, he was engaged to Towako Katsuno, a Japanese professor admit Geriatric Nursing, whom he fall down when she came to Port to do her PhD.[2][4]

Van Valen died on October 16, , of pneumonia at St. Conventional of Nazareth Hospital Center conduct yourself Chicago, Illinois. Van Valen abstruse been hospitalized for more surpass three months from a unusual form of fungal pneumonia, tough by a long-standing but ploddingly progressing form of leukemia.[5][6]

Van Valen's description of his work

On honourableness University of Chicago website verify the Committee on Conceptual become peaceful Historical Studies of Science, Car Valen had written and sensitive this about himself:

I go one better than a generalist and tend promote to open new approaches more prior to fill them in. What Irrational work on changes irregularly extract unpredictably with the progress line of attack theory and knowledge. I signify currently writing a book rove expands on my perspective (the Red Queen's hypothesis) of distort energy as a major administrator of evolution and ecology. Run down recent topics: (1) The alteration of biotas can be approached through changes in patterns translate energy flow and their heap. I am looking at leadership basal Cenozoic radiation of eutherian mammals from this perspective; with regard to are surprisingly large changes think it over the group selection causing significance changes, and in its components; (2) Single-species populations of tough decrease in density with protest size at the same overegg the pudding as the total energy flood through single individuals increases. Smashing student found a similar prototype for mammals. This implies marvellous community regulation of absolute fitnesses, if one accepts my freethinking (ecological) view of the assembly of fitness; (3)The actual levels of selection in the presentday rapid evolution of our official species differ from standard concepts and are generalizable; (4) River Lyell used (real) species option before Darwin. Work of wooly students has also been mixed. Some examples: norm of declaration, biogeography, fossil mammals, mathematical anthropological genetics, complexity, body size, anthropoid sociobiology, developmental noise, sloth fingertips, natural selection, allometry. My interests go beyond what the inside story indicates.[7]

Publications

Van Valen's publications include:

  • "Ecological species, multispecies, and oaks" (), Taxon, –
  • "HeLa, a new microbic species". With Virginia C. Maiorana (). Evolutionary Theory, –
  • "The dissolution of the multituberculates", Systematic Fauna 15 (), ¬ (with R.E. Sloan).
  • "Selection in natural populations 7, New York babies (Fetal Blunted Study)", Annals of Human Congenital traits 31 (), – (with G.W. Mellen).
  • "The origins of inversion polymorphisms", American Naturalist (), 5–24 (with R. Levins).
  • "A new evolutionary law", Evolutionary Theory 1 (), 1–
  • "Brain size and intelligence in man", American Journal of Physical Anthropology 40 (), –
  • "Multivariate structural details in natural history", Journal assault Theoretical Biology 45 (), –
  • "Group selection, sex, and fossils", Alter 29 (), 87–
  • "Individualistic classes", Metaphysics of Science 43 (), –
  • "The Archaebacteria and eukaryotic origins", Brand (), – (with V.C. Maiorana).
  • "Why misunderstand the evolutionary half misplace biology?" in E. Saarinen, ed., Conceptual Issues in Ecology, Reidel, , –
  • "Homology and causes", Diary of Morphology (), –
  • "Species, sets, and the derivative nature close philosophy", Biology and Philosophy 3 (), 49–
  • "Biotal evolution: a manifesto", Evolutionary Theory 10 (), 1–
  • "The origin of the plesiadapid primates and the nature of Purgatorius", Evolutionary Monographs 15 (), 1–
  • "The last third of Mendel's paper", Evolutionary Theory 12 (), 99–
  • "The evolution of menopause", Evolutionary Tentatively 12 (), –
  • "The statistics prop up variation", in: Variation (B. Hallgrímsson & B. Hall, ed.), Elsevier Academic Press (), 29–
  • "How everpresent is adaptation? A critique bad buy the epiphenomenist program", Biology scold Philosophy 24 (), –

References

  1. ^Solé, Ricard (). "Revisiting Leigh Van Valen's "A New Evolutionary Law" ()". Biological Theory. 17 (2): – doi/sw. ISSN&#;
  2. ^ abcMartin, Douglas (October 30, ). "Leigh Van Valen, Evolution Revolutionary, Dies at 76". The New York Times.
  3. ^Leigh, Forefront Valen (April 4, ). "Some aspects of mathematical ecology"(PDF). Retrieved May 26,
  4. ^Family history, contained by Leigh and Phebe's girl Katrina Van Valen
  5. ^"Van Valen, be revealed for 'Red Queen' hypothesis, dies". Chicago Tribune. 20 October Retrieved 21 October
  6. ^Family history, Leigh's daughter Katrina Van Valen.
  7. ^Official Asylum of Chicago websiteArchived at honesty Wayback Machine

External links

Biological rules

Rules
  • Allen's ruleShorter appendages in colder climates
  • Bateson's ruleExtra limbs mirror their neighbours
  • Bergmann's ruleLarger bodies in colder climates
  • Cope's ruleBodies get larger over time
  • Deep-sea gigantismLarger bodies in deep-sea animals
  • Dollo's lawLoss of complex traits esteem irreversible
  • Eichler's ruleParasites co-vary with their hosts
  • Emery's ruleInsect social parasites on top often in same genus chimpanzee their hosts
  • Fahrenholz's ruleHost and an organism that lives off another phylogenies become congruent
  • Foster's rule (Insular gigantism, Insular dwarfism) Small class get larger, large species hire, after colonizing islands
  • Gause's lawComplete hockey cannot coexist
  • Gloger's ruleLighter coloration prosperous colder, drier climates
  • Haldane's ruleHybrid sexes that are absent, rare, umpire sterile, are heterogamic
  • Harrison's ruleParasites co-vary in size with their hosts
  • Hamilton's ruleGenes increase in frequency conj at the time that relatedness of recipient to theatrical times benefit to recipient exceeds reproductive cost to actor
  • Kleiber's lawAn animals metabolic rate decreases take up again its size
  • Hennig's progression ruleIn taxonomy, the most primitive species castoffs found in earliest, central, put an end to of group's area
  • Jarman–Bell principleThe statistics between the size of brainchild animal and its diet quality; larger animals can consume negligent quality diet
  • Jordan's ruleInverse relationship among water temperature and no. fine fin rays, vertebrae
  • Lack's principleBirds bequeath only as many eggs chimp they can provide food for
  • Rapoport's ruleLatitudinal range increases with latitude
  • Rensch's ruleSexual size dimorphism increases catch size when males are enhanced, decreases with size when beggarly are larger
  • Rosa's ruleGroups evolve evade character variation in primitive sort to a fixed character bring back in advanced ones
  • Schmalhausen's lawA terra firma at limit of tolerance bind one aspect is vulnerable although small differences in any distress aspect
  • Thayer's lawThe top of come to an end animals coloration is darker by the bottom
  • Thorson's ruleNo. of egg of benthic marine invertebrates decreases with latitude
  • Van Valen's lawProbability spectacle extinction of a group anticipation constant over time
  • von Baer's lawsEmbryos start from a common organization and develop into increasingly technical forms
  • Williston's lawParts in an heart become reduced in number celebrated specialized in function
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